Pablo Marquet
Departamento de Ecología
Pontificia Universidad Católica de Chile
Casilla 114-D, Santiago
Correo electrónico: pmarquet@bio.puc.cl
www.bio.puc.cl/profs/marquet
Teléfono: 56-2-3542639
Fax: 56-2-6862621

Research:
My first research interests lie in the areas of macroecology and ecological complexity. The main focus here is the search for the general principles that underlie the seemingly endless complexity, diversity and variability of ecological systems. Because of its synthetic character, our research is neither restricted to a particular temporal or spatial scale of analysis, nor limited to a particular level of ecological organization, thus encompassing phenomena in both ecological and evolutionary time scales and from local communities to continental and global biotas. What matters is the question and way ecological systems are approached. As in other branches of science, the search for general principles and invariants in complex systems takes the form of statistical regularities such as scaling relationships and power law distributions in ecological attributes. Our challenge is to explain their mechanistic basis and make predictions of unknown relationships. My second interest is in ecoinformatics and conservation biogeography. This research is focused on trying to apply ecological knowledge and available large-scale datasets to help inform conservation policy and management. Our main aim is to help provide the knowledge base needed to effectively preserve Chilean biodiversity through conservation planning tools associated with the design of protected area networks, gap analysis, niche modeling and analysis of biogeographic patterns than can help to understand the mechanisms behind the generation and maintenance of biodiversity, and the way it will be affected by global change.

Education:
University of Concepción, Licenciatura en Biología, 1987.
University of New Mexico, Ph.D. (Biology), 1993.
Catholic University of Chile, Postdoctoral fellow, 1994.

Present academic positions:
Full Professor, Catholic University of Chile.
Associate Investigator, Institute of Ecology and Biodiversity (
IEB).
Principal Investigator, Center for Advanced Studies in Ecology and Biodiversity (CASEB).

Honors and prizes:
Graduate Student Achievement Award, University of New Mexico, 1993.
Outstanding Young Scientist Award, Biological Society of Chile, 1995.
Outstanding Paper in the Discipline of Landscape Ecology Award, U.S. Chapter of the International Association for Landscape Ecology.
"Frontiers of Science", Program, Chilean Academy of Sciences, 2004-2006.
Member of the Faculty of 1000.

  Publications 
 -1_Kelt_etal_2016_Jour_Arid_Envirom.pdf

Kelt, D.A., H. Cofre, C. Cornelius,  A. Engilis Jr., J.R. Gutiérrez, P.A. Marquet, R. Medel, P.L. Meserve, V. Quirici, H. Samaniego & R.A. Vásquez (2016) The avifauna of Bosque Fray Jorge National Park and Chile´s Norte Chico. Journal of Arid Environments 126: 23-36.

 -1_Castaneda_et_al_2015_EcolEvol.pdf

Castaneda, L.E., K. Godoy, M. Manzano, P. A. Marquet & O. Barbosa (2015) Comparison of soil microbial communities inhabiting vineyards and native sclerophyllous forests in central Chile. Ecology and Evolution 5 (18): 3857–3868.

 -1_Xu_etal_2015_Eco_Appli.pdf

Chi Xu, M. Holmgren, E.H. Van Nes, F. T. Maestre, S. Soliveres, M. Berdugo, S. Kefi, P. A. Marquet, S.Abades, & M. Scheffer (2015) Can we infer plant facilitation from remote sensing? a test across global drylands. Ecological Applications, 25(6): 1456–1462.

 -1_Badano_etal_2015_Jour_Veg_Scien.pdf

Badano, E,I.,  R.O Bustamante, E. Villarroel, P. A. Marquet & L.A.  Cavieres (2015) Facilitation by nurse plants regulates community invasibility in harsh environments.  Journal of Vegetation Science 26: 756–767.

 -1_Marquet_etal_2015_BioScience.pdf

Marquet PA, AP Allen,  JH Brown, JA Dunne, BJ Enquist, JF Gillooly, PA Gowaty, J Harte, SP Hubbell, JG. Okie, A Ostling,  M Ritchie, D Storch, and GB West (2015) On the importance of  first principles in ecological theory development. BioScience 65: 342-343.

 

Scheffer, M., J. Bascompte, T. K. Bjordam, S. R. Carpenter, L. B. Clarke, C. Folke, P.A Marquet, N. Mazzeo, M. Meerhoff, O. Sala, and F. R. Westley (2015) Dual thinking for scientists. Ecology and Society 20(2): 3.

 -1_Xu_etal_2015_Eco_Appli.pdf

Xu,C., M., Holmgren, E.H. Van Nes, F. T. Maestre, S. Soliveres, M. Berdugo, S. Kefi, P. A. Marquet, S.Abades, & M. Scheffer (2015) Can we infer plant facilitation from remote sensing? a test across global drylands. Ecological Applications 25:  1456–1462.

 -1_Abades_et_al-2014-Journal_of_Ecology.pdf

Abades, S. R., A. Gaxiola & P.A. Marquet (2014) Fire, percolation thresholds and the savanna forest transition: a neutral model approach. Journal of Ecology 102: 1386 – 1393.

 -1_Borthagaray_etal_2014_PRS_B.pdf

Borthagaray, A. I., Arim, M., & P. A. Marquet (2014) Inferring species roles in metacommunity structure from species co-occurrence networks. Proceedings of the Royal Society B: Biological Sciences 281(1792), 2014.1425.

 -1_Marquet_etal_2014_BioScience.pdf

Marquet, P. A., A. P Allen, J. H. Brown, J. A. Dunne, B.J. Enquist, J. F. Gillooly, P. A. Gowaty, J. L. Green, J. Harte, S. P. Hubbell, J. O´Dwyer, J. G Okie, A. Ostling, M. Ritchie, D. Storch & G. B West  (2014) On theory in ecology. BioScience 64: 701-710.

 -1_Nuñez-Lopez_etal_2014_Discret_Cont_Dynam.pdf

Nunez-Lopez M., J.X. Velasco-Hernandez & P.A. Marquet (2014) The dynamics of techniological change under constraints: Adopters and resourcers. Discrete and Continuous Dynamical Systems Series B 19: 3299-3317.

 -1_Patron_etal_2014_Mastozoologia.pdf

Patron, H. Z., E. Palma, P. A. Marquet, & G. Ceballos (2014) Phylogenetic relationships of calomys sorellus complex (rodentia: cricetidae), with the description of two new species. Revista Mexicana de Mastozoología 4: 1-23.

 

Keymer J.E. & P. A. Marquet (2014) The complexity of cancer ecosystems, In: Frontiers in Ecology, Evolution and Complexity by Mariana Benítez, Octavio Miramontes & Alfonso Valiente-Banuet ISBN: 978-1-938128-05-9.

 

Hannah, L., Roehrdanz, P. R., Ikegami, M., Shepard, A. V., Shaw, M. R., Tabor, G., P.A. Marquet & Hijmans, R. J. (2013) Climate change, wine, and conservation. Proceedings of the National Academy of Sciences 110(17):6907-6912.

 -1_Samaniego_Marquet_2013_Theor_Ecol.pdf

Samaniego, H &  P. A. Marquet (2013) Range structure analysis: unveiling the internal structure of species’ ranges. Theoretical Ecology 6:419–426.

 -1_Araujo_etal_2013_Ecol_Lett.pdf

Araujo, M.B., F. Ferri-Yanez, F. Bozinovic, P.A. Marquet, F. Valladares & S.L. Chown (2013) Heat freezes niche evolution. Ecology Letters 16:1206-1219.

 -1_Duran_etal_2013_PLOSONE.pdf

Duran, A. P., S. Casalegno, P.Marquet & K.Gaston (2013) Representation of Ecosystem Services by Terrestrial Protected Areas:Chile as a Case Study. PLoS ONE 8(12): e82643

 -1_González-Salazar_etal_2013_Ecolog_Modelling.pdf

Gonzales-Salazar, C., C.R. Stevens & P.A. Marquet (2013) Comparing the relative contributions of biotic and abiotic factors as mediators of species’ distributions. Ecological Modelling 248: 57– 70.

 -1_Hannah_etal_2013_PNAS_Reply.pdf

Hannah, L., P.R. Roehrdanz, M. Ikegami, A. V. Shepard, M. R. Shaw, G. Tabor, L. Zhi, P. A. Marquet & R. J. Hijmans (2013b) Reply to van Leeuwen et al.: Planning for agricultural adaptation to climate change and its consequences for conservation. Proceedings of the National Academy of Sciences 110: E3053.

 

Hernandez, C.,  E. Rodriguez-Serrano, J. Avaria-Llautureo, O. Inostroza-Michael, B. Morales-Pallero, D. Boric-Bargetto, C. Canales-Aguirre, P. A.  Marquet & A. Meade (2013) Using phylogenetic information and the comparative method to evaluate hypotheses in macroecology. Methods in Ecology and Evolution 4: 401-415.

 -1_Borthagaray_etal_2012_Oikos.pdf

Borthagaray, A.I., M. Arim & P. A. Marquet (2012) Connecting landscape structure and patterns in body size distributions. Oikos 121:697-710.

 -1_Barnosky_etal_2012_Nature.pdf

Barnosky, A.D., E. A. Hadly, J. Bascompte, E. L. Berlow, J. H. Brown, M. Fortelius, W. M. Getz, J. Harte, A. Hastings, P. A. Marquet,  N. D. Martinez, A. Mooers,  P. Roopnarine, G. Vermeij, J.W. Williams, R. Gillespie, J. Kitzes, C. Marshall, N. Matzke, D.P. Mindell, E. Revilla & A. B. Smith (2012) Approaching a state shift in Earth’s biosphere. Nature 486: 52–58

 -1_Shaw_etal_2012_Jour_Mammalogy.pdf

Shaw, A. K., J.L Galaz & P. A. Marquet (2012) Population dynamics of the vicuña (Vicugna vicugna): density-dependence, rainfall, and spatial distribution. Journal of Mammalogy 93(3):658–666.

 -1_Keymer_etal_2012_Theor_Ecol.pdf

Keymer, J.E., M.A. Fuentes & P.A. Marquet (2012) Diversity emerging: from competitive exclusion to neutral coexistence in ecosystems. Theoretical Ecology 5 (3):457-463.

 -1_Marquet_etal_2012_PNAS.pdf

Marquet, P., C. M. Santoro, C. Latorre, V. G. Standen, S. R. Avades, M. M. Rivadeneira, B. Arriaza & M. E. Hochberg (2012) Emergence of social complexity among coastal hunter-gatherers in the Atacama Desert of northern Chile. PNAS 109(37):14754-114760.

 -1_Mulder_etal_2012_Advan_Ecol_Research.pdf

Mulder, C., A. Boit, S. Mori, J.A. Vonk, S.D. Dyer, L. Faggiano, S. Geisen, A. L. González, M. Kaspari, S. Lavorel, P. A. Marquet, A. G. Rossberg, R. W. Sterner, W. Voigt & D. H. Wall (2012) Distributional (In)Congruence of Biodiversity-Ecosystem functioning. Advances in Ecological Research 46: 1-88.

 Abades_Marquet_2011_Biol_Res.pdf

Abades, S.R & P.A. Marquet (2011) Finite size scaling in the local abundances ofgeographic populations. Biological Research 44(1):107-112.

 Gonzalez_etal_2011_Oikos.pdf

González, A.L.,  J.M. Fariña, A.D. Kay, R.Pinto & P.A. Marquet (2011) Exploring patterns and mechanisms of interspecific and intraspecific variation in body elemental composition of desert consumers.  Oikos 120:1247-1255.

 Gonzalez_etal_2011_Oecologia.pdf

González, A.L., J.M. Fariña, R. Pinto, C. Pérez, K.C. Weathers, J. Armesto & P. Marquet (2011) Bromeliad growth and stoichiometry: responses to atmosphericb nutrient supply in fog-dependent ecosystems of the hyper-arid Atacama Desert, Chile. Oecología 167:835–845

 27_Morlon_et_al_2011_Ecology_letters.pdf

Morlon, H., D.W. Schwilk, J.A. Bryant, P.A. Marquet, A.G. Rebelo, C. Tauss, B.J. M. Bohannan & J. L. Green (2011) Spatial patterns of phylogenetic diversity. Ecology Letters 14: 141-149

 

Núñez- Ávila, M. C., M. Uriarte, P.A. Marquet & J.J Armesto (2011) Microsatellite markers for the relict tree Aextoxicon punctatum: the only species in the chilean endemic family Aextoxicaceae. American Journal of Botany 98 (2):e30–e32

 Arim_etal_2010_Oikos.pdf

Arim, M., S.R. Abades, G. Laufer, M. Loureiro & P.A. Marquet (2010) Food web structure and body size: trophic position and resource acquisition. Oikos 119: 147-153.

 Armesto_etal_2010_LUP.pdf

Armesto, J.J., D. Manuscevich, A. Mora., C. Smith-Ramírez, R. Rozzi, A.M. Abarzúa & P.A. Marquet (2010) From the Holocene to the Anthropocene: A historical framework for land cover change in southwestern South America in the past 15,000 years. Land Use Policy 27:148-160.

 Badano_etal_2010_Act_Oecol.pdf

Badano, E.I., P.A. Marquet & L.A. Cavieres (2010) Predicting effects of ecosystem engineering on species richness along primary productivity gradients. Acta Oecologica 36: 46-54.

 Barbosa_etal_2010_Funct_Ecol.pdf

Barbosa, O., P.A. Marquet, L.D. Bacigalupe, D.A. Christie, E. del-Val, A.G. Gutierrez,  C.G. Jones, K.C. Weathers & J.J. Armesto (2010) Interactions among patch area, forest structure and water fluxes in a fog-inundated forest ecosystem in semi-arid Chile. Functional Ecology 24: 909-917.

 Borthagaray_etal_2010_Theor_Biol.pdf

Borthagaray, A.I., M.A. Fuentes & P.A. Marquet (2010) Vegetation pattern formation in a fog-dependent ecosystem. Journal of Theoretical Biology 265: 18-26.

 Rivadeneira_etal_2010_Aqua_Conserv.pdf

Rivadaneira, M.M., C.M. Santoro & P.A. Marquet (2010) Reconstructing the history of human impacts on coastal biodiversity in chile: constraints and opportunities. Aquatic Conservation: Marine and freshwater ecosystems. Aquatic Conservation: Marine and Freshwater Ecosystems 20: 74-82.

 Aldana_et_al_2009_Jour_Parasit.pdf

Aldana, M., K. Gonzalez, G. Loot, J. Pulgar & P.A. Marquet (2009) First intermediate host of the digenean trematode Proctoeces lintoni (Fellodistomidae) in Chile. Journal of Parasitology 95: 1408-1414.

 Badano_Marquet_2009_Plant_Ecol_Evol_Syst.pdf

Badano, E.I. & P.A. Marquet (2009) Biogenic habitat creation affects biomass-diversity relationships in plant communities. Perspectives in Plant Ecology, Evolution and Systematics 11: 191-201.

 Fernandez_etal_2009_Ecol_Lett.pdf

Fernández, M., A. Astorga, S.A. Navarrete, C. Valdovinos & P.A. Marquet (2009) Deconstructing latitudinal species richness patterns in the ocean: does larval development hold the clue?. Ecology Letters 12: 601-611.

 Marquet_2009_revista_chilena.pdf

Marquet, P.A. (2009) Beyond Darwin: On the role of niche construction and self-organization in evolution. Revista Chilena de Historia Natural 82: 493-496.

 Marquet_2009_Princeton_Guide_to_Ecology_Chapter.pdf

Marquet, P.A. (2009) Macroecological approaches to communities and ecosystems. In (S.A. Levin) Princeton Guide to Ecology 386-394. Princeton University Press, USA.

 

Parra, L.E., R. Villagran-Mella & P.A. Marquet (2009) Phylogeny of the genera Euclidiodes and Hasodima (Lepidoptera: Geometridae) and description of two new species from the Fray Jorge relict forest in northern Chile. Zootaxa 2273: 59-68.

 -1_Ramos_etal_2009_Oecologia.pdf

Ramos-Jiliberto, R., A.A. Albornoz, F.S. Valdovinos, C. Smith-Ramírez, M. Arim, J.J. Armesto & P.A. Marquet (2009) A network analysis of plant-pollinator interactions in southern rain forests of Chiloé Island, Chile. Oecologia 160: 697-706.

 -1_Samaniego_Marquet_2009_RCHN.pdf

Samaniego, H. & P.A. Marquet (2009) Mammal and butterfly species richness in Chile: taxonomic covariation and history. Revista Chilena de Historia Natural 82: 135-151.

 -1_Tognelli_etal_2009_Biol_Cons.pdf

Tognelli, M.F., M. Fernández & P.A. Marquet (2009) Assessing the performance of the existing and proposed network of marine protected areas to conserve marine biodiversity in Chile. Conservation Biology 142: 3147-3153.

 

Ramírez de Arellano, P.I., M.F. Tognelli, C. Garín & P.A. Marquet (2008) Vacíos de Conservación y Sitios Prioritarios para la Conservación de los Vertebrados Nativos de la Región de Atacama. In: Libro Rojo de la Flora Nativa y de los Sitios Prioritarios para su conservación: Región de Atacama (F.A. Squeo, G. Arancio &  J.R. Gutiérrez, Eds.). Pp. 251-266, Ediciones Universidad de La Serena, La Serena, Chile. 

 Badano_Marquet_2008_Oecologia.pdf

Badano, E.I. & P.A. Marquet (2008) Ecosystem engineering affects ecosystem functioning in high-Andean landscapes. Oecologia 155: 821-829.    

 Gutierrez_etal_2008_Glob_Chang_Biol.pdf

Gutiérrez, A.G., O. Barbosa, D.A. Christie, E. del Val, H.A. Ewing, C.G. Jones, P.A. Marquet, K.C. Weathers & J.J. Armesto. (2008) Regeneration patterns and persistence of the fog-dependent Fray Jorge forest in semiarid Chile during the past two centuries. Global Change Biology 14: 161-176.

 Lafferty_etal_2008_Ecol_Lett.pdf

Lafferty, K.D., S. Allesina, M. Arim, C.J. Briggs, G. De Leo, A.P. Dobson, J.A. Dunne, P.T.J. Johnson, A.M. Kuris, D.J. Marcogliese, N.D. Martinez, J. Memmott, P.A. Marquet, J.P. McLaughlin, E.A. Mordecai, M. Pascual,  R. Poulin & D.W. Thieltges (2008) Parasites in food webs: the ultimate missing links. Ecology Letters  11: 533-546.

 Lenoir_etal_2008_Science.pdf

Lenoir, J., J.C. Gégout, P.A. Marquet, P. de Ruffray & H. Brisse (2008). A significant upward shift in plant species optimum elevation during the 20th century. Science 320: 1768-1771.

 Napolitano_etal_2008_Molec_Ecol.pdf

Napolitano, C., M. Bennett, W.E. Johnson, S.J. O'Brien, P.A. Marquet, I. Barria, E. Poulin & A. Iriarte (2008) Ecological and biogeographical inferences on two sympatric and enigmatic Andean cat species using genetic identification of faecal samples. Molecular Ecology 17: 678-690.    

 Tognelli_etal_2008_Divers_Distrib.pdf

Tognelli, M.F., P.I. Ramirez de Arellano & P.A. Marquet (2008) How well do the existing and proposed reserve networks represent vertebrate species in Chile? Diversity and Distributions 14: 148–158.

 -1_Arim_etal_2007_Oikos.pdf

Arim, M., F. Bozinovic & P.A. Marquet (2007) On the relationship between trophic position, body mass and temperature: reformulating the energy limitation hypothesis. Oikos 116: 1524-1530.

 -1_Arim_etal_2007_Amer_Natur.pdf

Arim, M., P.A. Marquet & F.M. Jaksic (2007) On the relationship between productivity and food chain length at different ecological levels. The American Naturalist 169: 62-72.

 -1_Badano_etal_2007_Jour_Ecol.pdf

Badano, E.I., E. Villarroel, R.O. Bustamante, P.A. Marquet & L.A. Cavieres (2007) Ecosystem engineering facilitates invasions by exotic plants in high-Andean ecosystems. Journal of Ecology 95: 682-688.

 -1_Cofre_etal_2007_Divers_Distrib.pdf

Cofré, H.L., K. Böhning-Gaese & P.A. Marquet (2007) Rarity in Chilean forest birds: Which ecological and life-history traits matter? Diversity and Distribution 13: 203-212.

 -1_Cofre_etal_2007_Quintessential.pdf

Cofré, H.L., H. Samaniego & P.A. Marquet (2007) Patterns of small mammal species richness in Mediterranean and Temperate Chile. In: (D.A. Kelt, E.P. Lessa,  J.A. Salazar-Bravo & J.L. Patton, eds.), “The Quintessential Naturalist: Honoring the Life and Legacy of Oliver P. Pearson”, pp. 275-302. University of California Publications in Zoology, University of California Press, California.

 -1_delVal_etal_2007_Oecologia.pdf

del-Val, E., J.J. Armesto, O. Barbosa & P.A. Marquet (2007) Effects of herbivory and patch size on tree seedling survivorship in a fog-dependent coastal rainforest in semiarid Chile. Oecologia 153: 625-632.

 -1_Labra_etal_ 2007_PNAS.pdf

Labra, F.A., P.A. Marquet & F. Bozinovic (2007) Scaling metabolic rate fluctuations. Proceedings on the National Academy of Sciences (USA) 104: 10900-10903.

 -1_Marquet_etal_2007_Scaling_Biodiversity.pdf

Marquet, P.A., S.R. Abades & F.A. Labra (2007) Biodiversity power laws. In: (D. Storch, P.A. Marquet & J.H. Brown, eds), "Scaling Biodiversity". Cambridge University Press, Cambridge.

 

Marquet, P.A., S. Abades, J.E. Keymer & H. Zeballos (2007) Discontinuities in body size distributions: a view from the top. In: (C.R. Allen & C.S. Holling, eds), "Discontinuity Theory in Ecosystems and Other Complex Systems". Complexity Series, Columbia University Press, New York).

 -1_McGill_etal_2007_Ecol_Lett.pdf

McGill, B.J., R.S. Etienne, J.S. Gray, D. Alonso, M.J. Anderson, H.K.Benecha, M. Dornelas, B.J. Enquist, J.L. Green, F. He, A.H. Hurlbert, A.E. Magurran, P.A. Marquet, B.A. Maurer, A. Ostling, C.U. Soykan, K.I. Ugland & E.P. White (2007) Species abundance distributions: moving beyond single prediction theories to integration within an ecological framework. Ecology Letters 10: 995–1015.

 -1_Rivadeneira_Marquet_2007_Paleobiology.pdf

Rivadeneira, M.M. & P.A. Marquet (2007) Selective extinction of late Neogene bivalves on the temperate Pacific coast of South America. Paleobiology 33: 455-468.

 -1_Vergara__Marquet_2007_Land_Ecol.pdf

Vergara, P.M. & P.A. Marquet (2007) On the seasonal effect of landscape structure on a bird species: the thorn-tailed rayadito in a relict forest in northern Chile. Landscape Ecology 22: 1059-1071.

 -1_Wallem_etal_2007_RCHN.pdf

Wallem, P.K., C.G. Jones, P.A. Marquet & F.M. Jaksic (2007) Identificación de los mecanismos subyacentes a la invasión de Castor canadensis (Rodentia) en el archipiélago de Tierra del Fuego, Chile. Revista Chilena de Historia Natural 80: 309-325.

 -1_Wilson_etal_2007_PLoS_Biology.pdf

Wilson, K.A., E.C. Underwood, S.A. Morrison, K.R. Klausmeyer, W.W. Murdoch, B. Reyers, G. Wardell-Johnson, P.A. Marquet, P.W. Rundel, M.F. McBride, R.L. Pressey, M. Bode, J.M. Hoekstra, S.J. Andelman, M. Looker, C. Rondinini, P. Kareiva, M.R. Shaw & H.P. Possingham (2007) Conserving biodiversity efficiently: What to do, where and when. PLoS Biology 5: 1850-1861.

 -1_Allen_etal_2006_Ecol_Lett.pdf

Allen, C.R., A.S. Garmestani, T.D. Havlicek, P.A. Marquet, G.D. Peterson, C. Restrepo, C.A. Stow & B.A. Weeks (2006) Patterns in body mass distributions: Sifting among alternative hypotheses. Ecology Letters 9: 630-643.

 -1_Arim_etal_2006_PNAS.pdf

Arim, M., S.R. Abades, P.E. Neill, M. Lima & P.A. Marquet (2006) Spread dynamics of invasive species. Proceedings on the National Academy of Sciences (USA) 103: 374-378.

 -1_Arroyo_etal_2006_Chapter14_Diagnostico.pdf

Arroyo, M.T.K., J.J. Armesto, F. Bozinovic, L.A. Cavieres, J.R. Gutiérrez, C. Latorre, P. Marquet, D. Soto & F.A. Squeo (2006) Ciencias ambientales. Diagnóstico y mirada hacia el futuro. In: (J. Allende, J. Babul, S. Martínez & T. Ureta, eds), "Análisis y Proyecciones de la Ciencia Chilena 2005", pp. 295-331. Academia Chilena de Ciencias, Santiago.

 -1_Arroyo_etal_2006_BiodivChile.pdf

Arroyo, M.T.K., P.A. Marquet, C. Marticorena, J.A. Simonetti, L.A. Cavieres, F.A. Squeo, R. Rozzi & F. Massardo (2006) El Hotspot chileno, prioridad mundial para la conservación. In: (P. Saball, M.T.K. Arroyo, J.C. Castilla, C. Estades, J.M. Ladrón De Guevara, S. Larraín, C. Moreno, F. Rivas, J. Rovira, A. Sánchez &  L. Sierralta, eds), "Biodiversidad de Chile. Patrimonio y Desafíos", pp. 94-99. Comisión Nacional del Medio Ambiente. Santiago.

 -1_delVal_etal_2006_Ecosystems.pdf

del-Val, E., J.J. Armesto, O. Barbosa, D.A. Christie, A.G. Gutiérrez, C.G. Jones, P.A. Marquet & K.C. Weathers (2006) Rain forest islands in the Chilean semiarid region: fog-dependency, ecosystem persistence and tree regeneration. Ecosystems 9: 598-608.

 -1_MenaLorca_etal_2006_Mathe_Bioscience.pdf

Mena-Lorca, J., J.X. Velasco-Hernandez & P.A. Marquet. (2006) Coexistence in metacommunities: a three-species model. Mathematical Biosciences 202: 42-56.

 -1_Pinto_etal_2006_Jour_Arid_Env.pdf

Pinto, R., I. Barría & P.A. Marquet (2006) Geographical distribution of Tillandsia lomas in the Atacama desert, northern Chile. Journal of Arid Environments 65: 543-552.

 -1_Cornelius_etal_2005_Bosques_Costeros.pdf

Cornelius, C., S. Reid & P.A. Marquet (2005) Conservación de la avifauna de los bosques relictos del Norte Chico y su relación con los bosques costeros valdivianos. In: (C. Smith-Ramírez, J.J. Armesto& C. Valdovinos, eds), "Historia, Biodiversidad y Ecología de los Bosques Costeros de Chile", pp. 134-138. Editorial Universitaria, Santiago.

 -1_Hidalgo_etal_2005_PhysicaA.pdf

Hidalgo, C.A., F. Claro & P.A. Marquet (2005) Stationary state structure of a random copying mechanism over a complex network. Physica A 353: 674-684.

 -1_Labra_etal_2005_Species_Invasions.pdf

Labra, F.A., S.R. Abades & P.A. Marquet (2005) Distribution and abundance. Scaling patterns in exotic and native bird species. In: (D.F. Sax, J.J. Stachowicz & S.D. Gaines, eds), "Species Invasions. Insights into Ecology, Evolution and Biogeography", pp. 421-446. Sinauer Associates Inc, Massachusetts.

 -1_Marquet_etal_2005_Jour_Exp_Biol.pdf

Marquet, P.A., R.A. Quiñones, S. Abades, F. Labra, M. Tognelli, M. Arim & M. Rivadeneira (2005) Scaling and power-laws in ecological systems. Journal of Experimental Biology 208: 1749-1769.

 -1_Palma_etal_2005_Jour_Biogeogr.pdf

Palma, R.E., P.A. Marquet & D. Boric-Bargetto (2005) Inter- and intraspecific phylogeography of small mammals in the Atacama Desert and adjacent areas of northern Chile. Journal of Biogeography 32: 1931-1941.

 -1_Palma_etal_2005_Jour_Mammal.pdf

Palma, R.E., E. Rivera-Milla, J. Salazar-Bravo, F. Torres-Pérez, U.F.J. Pardiñas, P.A. Marquet, A. Spotorno, A.P. Meynard & T.L. Yates (2005) Phylogeography of Oligoryzomys longicaudatus (Rodentia: Sigmodontinae) in temperate South America. Journal of Mammalogy 86: 191-200

 -1_Santoro_etal_2005_Global_desert_people.pdf

Santoro, C.M., B.T. Arriaza, V.G. Standen & P.A. Marquet (2005)  People of the coastal Atacama desert: living between sand dunes and waves of the Pacific Ocean. In: (P. Veth, M. Smith & P. Hiscock, eds), "Desert Peoples: Archaeological Perspectives", pp. 243-260. Blackwell Publishing, Oxford.

 -1_Santoro_etal_2005_23S.pdf

Santoro, C.M., V.G. Standen, B.T. Arriaza & P.A. Marquet (2005) Hunter-gatherers on the coast and hinterland of the Atacama Desert. In: (M. Smith & P. Hesse, eds), "23º South: The Archaeology and Environmental History of the Southern Desert", pp. 172-185. National Museum of Australia Press, Canberra.

 -1_Storch_etal_2005_Science.pdf

Storch, D., P.A. Marquet & K.J. Gaston (2005) Untangling an entangled bank. Science 307: 684-686.

 -1_Tognelli_etal_2005_Biol_Conser.pdf

Tognelli, M.F., C. Silva-García, F.A. Labra & P.A. Marquet (2005) Priority areas for the conservation of coastal marine vertebrates in Chile. Biological Conservation 126: 420-428.

Arim, M. & P.A. Marquet (2004) Intraguild predation: a widespread interaction related to species biology. Ecology Letters 7: 557-564.
Arroyo, M.T.K., P. Marquet, C. Marticorena, J. Simonetti, L.A. Cavieres, F.A. Squeo & R. Rozzi (2004) Chilean winter rainfall - Valdivian forests. In: (R.A. Mittermeier, P. Robles, M. Hoffmann, J. Pilgrim, T. Brooks, C. Goettsch-Mittermeier, J. Lamoreux & G.A.B. da Fonseca, eds), "Hotspots Revisited: Earth's Biologically Richest and Most Endangered Terrestrial Ecoregions",  pp. 99-103. CEMEX, México.
 Figueroa, J.A., S.A. Castro, P.A. Marquet & F.M. Jaksic. (2004) Exotic plant invasions to the mediterranean region of Chile: causes, history and impacts. Revista Chilena de Historia Natural 77: 465-483.
George-Nascimento, M., G. Muñoz, P.A. Marquet & R. Poulin (2004) Testing the energetic equivalence rule with helminth endoparasites of vertebrates. Ecology Letters 7: 527-531.
Marquet, P.A. & G.A. Bradshaw (2004) Symposium: Ecosystem disruption in the Americas. Revista Chilena de Historia Natural 77: 369-370.

Marquet, P.A., M. Fernández, S.A. Navarrete & C. Valdovinos (2004) Diversity emerging: Towards a deconstruction of biodiversity patterns. In: (M. Lomolino & L.R. Heaney, eds), "Frontiers of Biogeography: New Directions in the Geography of Nature", pp. 192-209. Cambridge University Press, Cambridge.
Marquet, P.A., F.A. Labra & B.A. Maurer (2004) Metabolic ecology: linking individuals to ecosystems. Ecology 85: 1794-1796
Meynard, C.N., H. Samaniego & P.A. Marquet (2004) Biogeografía de aves rapaces de Chile. In: (A. Muñoz, J. Rau & J. Yáñez, eds.), “Aves Rapaces de Chile”, pp. 129-143. Ediciones CEA, Valdivia.
Rodrigues, A.S.L., H.R. Akçakaya, S.J. Andelman, M.I. Bakarr, L. Boitani, T.M. Brooks, J.S. Chanson, L.D.C. Fishpool, G.A.B. da Fonseca, K.J. Gaston, M. Hoffman, P.A. Marquet, J.D. Pilgrim, R.L. Pressey, J. Schipper, W. Sechrest, S.N. Stuart, L.G. Underhill, R.W. Waller, M.E.J. Watts & X. Yan (2004) Global gap analysis: priority regions for expanding the global protected-area network. Bioscience 54: 1092-1100.
Rodrigues, A.S.L., S.J. Andelman, M.I. Bakarr, L. Boitani, T.M. Brooks, R.M. Cowling, L.D.C. Fishpool, G.A.B. da Fonseca, K.J. Gaston, M. Hoffman, J. Long, P.A. Marquet, J.D. Pilgrim, R.L. Pressey, J. Schipper, W. Sechrest, S.N. Stuart, L.G. Underhill, R.W. Waller, M.E.J. Watts & X. Yan (2004) Effectiveness of the global protected area network in representing species diversity. Nature 428: 640-643.
Smith, F.A., J.H. Brown, J.P. Haskell, S.K. Lyons, J. Alroy, E.L. Charnov, T. Dayan, B.J. Enquist, M. S.K. Ernest, E.A. Hadly, K.E. Jones, D.M. Kaufman, P.A. Marquet, B.A. Maurer, K. Niklas, W.P. Porter, K.B. Tiffney & M.R. Willig (2004) Similarity of mammalian body size across the taxonomic hierarchy and across space and time. The American Naturalist 163: 672-691.
Torres-Pérez, F., J. Navarrete-Droguett, R. Aldunate, T.L. Yates, G.J. Mertz, P.A. Vial, M. Ferrés, P.A. Marquet & R.E. Palma (2004) Peridomestic small mammals associated with confirmed cases of human Hantavirus disease in southcentral Chile. American Journal of Tropical Medicine and Hygiene 70: 305-309.

Bradshaw, G.A. & P.A. Marquet (eds) (2003) "How Landscapes Change. Human Disturbance and Ecosystem Fragmentation in the Americas". Springer-Verlag, New York.
Labra, F., N. Lagos & P.A. Marquet (2003) Dispersal and transients dynamics in metapopulations. Ecology Letters 6: 197-204.
Marquet, P.A. & G.A. Bradshaw (2003) Human disturbance and ecosystem fragmentation in the Americas. Synthesis and final reflections. In: (G.A. Bradshaw & P.A. Marquet, eds), "How Landscapes Change. Human Disturbance and Ecosystem Fragmentation in the Americas", pp. 345-353 . Springer-Verlag, New York.
Marquet, P.A., J.E. Keymer & H. Cofré (2003) Breaking the stick in space : of niche models, metacommunities, and patterns in the relative abundance of species. In: (T.M. Blackburn & K.J. Gaston, eds), “Macroecology: Concepts and Consequences”, pp. 64-84. Blackwell Science, Oxford, UK.
Marquet, P.A., J.X. Velasco-Hernández & J.E. Keymer (2003) Patch dynamics, habitat degradation, and space in metapopulations. In: (G.A. Bradshaw & P.A. Marquet, eds), "How Landscapes Change. Human Disturbance and Ecosystem Fragmentation in the Americas", pp. 239-254 . Springer-Verlag, New York.
Smith, F.A., S.K. Lyons, S.K.M. Ernest, K.E. Jones, D.M. Kaufman, T. Dayan, P.A. Marquet, J.H. Brown & J.P. Haskell (2003) Body mass of late quaternary mammals. Ecology 84: 3403.
Valdovinos, C., S.A. Navarrete & P.A. Marquet (2003) Mollusk species diversity in the Southeastern Pacific: Why are there more species towards the pole? Ecography 26:139-144.
Barbosa, O. & P.A. Marquet (2002) Effects of forest fragmentation on the beetle assemblage at the relict forests of Fray Jorge, Chile. Oecologia 132: 296-306.
Marquet, P.A. (2002) The search for general principles in ecology. Nature 418: 723.
Marquet, P.A. (2002) Of predators, prey, and power laws. Science 295: 2229-2230.
Marquet, P.A. (2002) Metapopulations. In: (H.A. Mooney & J. G. Canadell, eds), "Encyclopedia of Global Environmental Change. Volume II. The Earth System: Biological and Ecological Dimensions of Global Environmental Change", pp. 411-420. John Wiley & Sons Ltd., Chichester, London.
Marquet, P.A. & M.C. Depassier (2002) Kleiber and Planck. The missing link? Response. Science 298: 60-61.

Marquet, P.A., H. González, R. Pinto, C. Santoro, V. Standen & H. Zeballos (2002) Coastal deserts of Perú and Chile. In: (R. Mittermeier, C.G. Mittermeier, P. Robles Gil & J. Pilgrim, eds.), "Wilderness: Earth's Last Wild Places", pp. 365-373. Chicago University Press, Chicago.
Palma, R.E., E. Rivera-Milla, T.L. Yates, P.A. Marquet & A.P. Meynard (2002) Phylogenetic and biogeographic relationships of the mouse opossum Thylamys (Didelphimorphia, Didelphidae) in southern South America. Molecular Phylogenetics and Evolution 25: 245-253.
Reid, S., C. Cornelius, O. Barbosa, C. Meynard, C. Silva-Garcia & P.A.  Marquet  (2002) Conservation of temperate forest birds in Chile: Implications from the study of an isolated forest relict. Biodiversity and Conservation 11: 1975-1990.
Alvarado, J.L., R. Pinto, P.A. Marquet, C. Pacheco, R. Guiñez & J.C. Castilla (2001) Patch recolonization by the tunicate Pyura praeputialis in the rocky intertidal at the Bay of Antofagasta, Chile: evidence for self-facilitation mechanisms. Marine Ecology Progress Series 224: 93-101.
Carothers, J.H., F.M. Jaksic & P.A. Marquet (2001) Altitudinal zonation among lizards of the genus Liolaemus: questions answered and unanswered questions. Revista Chilena de Historia Natural 74: 313-316.
Cornelius, C., S.A. Navarrete & P.A. Marquet (2001) Effects of human activity on the structure of coastal marine bird assemblages in central Chile. Conservation Biology 15: 1396-1404.
Marquet, P.A. (2001) Ecology goes macro. Nature 412: 481-482.
Cornelius, C., H. Cofré & P.A. Marquet (2000) Effects of habitat fragmentation on bird species in a relict temperate forest in semiarid Chile. Conservation Biology 14: 534-543.
Fernández, M., E. Jaramillo, P.A. Marquet, C.A. Moreno, S.A. Navarrete, F.P. Ojeda, C.R. Valdovinos & J.A. Vasquez (2000) Diversity, dynamics and biogeography of Chilean benthic nearshore ecosystems: an overview and guidelines for conservation. Revista Chilena de Historia Natural 73: 797-830.
Kelt, D.A., P.A. Marquet & J.H. Brown (2000) Geographical ecology of South American desert small mammals: consequences of observations at local and regional scales. Global Ecology and Biogeography 9: 219-223.
Keymer, J.E., P.A. Marquet, J.X. Velasco-Hernández & S.A. Levin (2000) Extinction thresholds and metapopulation persistence in dynamic landscapes. The American Naturalist 156: 478-494.
Marquet, P.A. (2000) Invariants, scaling laws and ecological complexity. Science 289: 1487-1488.
Arroyo, M.T.K., R. Rozzi, J.A. Simonetti, P. Marquet & M. Salaberry (1999) Central Chile. In: (R.A. Mittermeier, N. Myers, P. Robles Gil & C. Goettsch Mittermeier, eds), "Hotspots: Earth's Biologically Richest and Most Endangered Terrestrial Ecorregions", pp. 161-171. CEMEX, México.
Bozinovic, F., J.A. Lagos & P.A. Marquet (1999) Geographic energetics of the Andean mouse  Abrothrix andinus. Journal of Mammalogy 80 (1): 205-209.
Cofré, H. & P.A. Marquet (1999) Conservation status, rarity, and geographic priorities for conservation of Chilean mammals: an assessment. Biological Conservation 88: 53-68.
Hernández-Suárez, C.M., P.A. Marquet & J.X. Velasco-Hernández (1999) Threshold parameters and metapopulation persistence. Bulletin of Mathematical Biology 61: 341-353.
Jaksic, F.M., J.C. Torres-Mura, C. Cornelius & P.A. Marquet (1999) Small mammals of the Atacama Desert (Chile). Journal of Arid Environments 42: 129-135.
Lima, M., P.A. Marquet & F.M. Jaksic (1999) El Niño events, precipitation patterns, and rodents outbreaks are statistically associated in semiarid Chile. Ecography 22: 213-218.
Marquet, P.A. & H. Cofré (1999) Large temporal and spatial scales in the structure of mammalian assemblages in South America: A macroecological approach. Oikos 85: 299-309.

Carothers, J.H., P.A. Marquet & F.M. Jaksic (1998) Thermal ecology of a Liolaemus lizard assemblage along an Andean altitudinal gradient in Chile. Revista Chilena de Historia Natural 71: 39-50.

Gutiérrez, J.R., F. López-Cortés & P.A. Marquet. (1998) Vegetation in an altitudinal gradient along the Río Loa in the Atacama desert of northern Chile. Journal of Arid Environments 40: 383-399.
Keymer, J.E., P.A. Marquet & A.R. Johnson (1998) Pattern formation in a patch occupancy metapopulation model: A cellular automata approach. Journal of Theoretical Biology 194: 79-90.
Lima, M., P.A. Marquet & F.M. Jaksic (1998) Population extinction risks of three Neotropical small mammal species. Oecología 115: 120-126

Marquet, P.A., F. Bozinovic, G.A. Bradshaw, C.C. Cornelius, H. González, J.R. Gutiérrez, E.R. Hajek, J.A. Lagos,  F. López-Cortés, L. Núñez, E.F. Rosello, C. Santoro, H. Samaniego, V.G. Standen,  J.C. Torres-Mura & F.M. Jaksic (1998) Los ecosistemas del desierto de Atacama y área andina adyacente en el norte de Chile. Revista Chilena de Historia Natural 71: 593-617.
Marquet, P.A. & M.L. Taper (1998) On size and area: Patterns of mammalian body size extremes across landmasses. Evolutionary Ecology 12: 127-139.
Santelices, B. & P.A. Marquet (1998) Seaweeds, latitudinal diversity patterns, and Rapoport's  rule. Diversity and Distributions 4: 71-75.
  Spotorno, A.E., H. Cofré, G. Manríquez, Y. Vilina, P.A. Marquet & L.I. Walker (1998) Genoma intermedio en una especie de mamífero filotino Loxodontomys en Chile central. Revista Chilena de Historia Natural 71: 359-373.
  Carothers, J.H., S.F. Fox, P.A. Marquet P.A. &  F.M. Jaksic (1997) Thermal characteristics of ten Andean lizards of the Chilean genus Liolaemus. Revista Chilena de Historia Natural 70: 297-309.
  Jaksic, F.M., P.A. Marquet & H. González (1997) A source-sink patch occupancy metapopulation model. Revista Chilena de Historia Natural 70: 371-380.
  Jaksic, F.M., P.A. Marquet & H. González (1997) Una perspectiva ecológica sobre el uso del agua en el norte grande: la región de Tarapacá como estudio de caso. Estudios Públicos 68: 171-195.
  Marquet, P.A. & M. Fox (1997) Southern temperate biota and ecosystems: Past, present, and future. Trends in Ecology and Evolution 12: 294-295.
  Marquet, P.A. & J.X. Velasco-Hernández (1997) A source-sink patch occupancy metapopulation model. Revista Chilena de Historia Natural 70: 371-380.
Torres-Contreras, H., E. Silva-Aranguiz, P.A. Marquet, P.A. Camus & F.M. Jaksic (1997) Spatiotemporal variability of rodent subpopulations at a semiarid Neotropical locality. Journal of Mammalogy 78: 505-513.
Brown, J.H., M.L.Taper & P.A. Marquet (1996) Darwinian fitness and reproductive power: reply to Koslowski. The American Naturalist 147: 1092-1097.
  Carothers, J.H., F.M. Jaksic & P.A. Marquet (1996) A model for species distributions along a gradient: Liolaemus lizards as study systems. Revista Chilena de Historia Natural 69: 301-307.
Keitt, T.H. & P.A. Marquet (1996) The introduced Hawaiian avifauna reconsidered: Evidence for self-organized criticality?. Journal of Theoretical Biology 182: 161-167.
Kelt, D.A., J.H. Brown, E.J. Heske, P.A. Marquet, S.R. Morton, J.R.W. Reid, K.A. Rogovin, & G. Shenbrot (1996) Community structure of desert small mammals: comparisons across four continents. Ecology 77: 746-761.
Lima, M., P.A. Marquet & F.M. Jaksic (1996) Extinction and colonization processes in subpopulations of five neotropical small mammal species. Oecologia 107: 197-203.
Taper, M.L. & P.A. Marquet (1996) How do species really divide resources? The American Naturalist 147: 1072-1086.
Marquet, P.A., S.A. Navarrete & J.C. Castilla (1995)  Body size, population density, and the energetic equivalence rule. Journal of Animal Ecology 64: 325-332.
Marquet, P.A. (1994) Diversity of small mammals in the pacific coastal desert of Peru and Chile and in the adjacent Andean area: Biogeography and community structure. Australian Journal of Zoology 42: 527-542.
Moreno, P.I., C. Villagrán, P.A. Marquet & L.G. Marshall (1994) Quaternary paleobiogeography of northern and central Chile. Revista Chilena de Historia Natural 67: 487-502.
Brown, J.H., P.A. Marquet & M.L. Taper (1993) Evolution of body size: Consequences of an energetic definition of fitness. The American Naturalist 142: 573-584.
  Marquet, P.A. (1993) Ambiente en las áreas fronterizas del norte de Chile: La frontera Chileno-Peruana y Chileno-Boliviana. In: (E. Tréllez, ed), "Ambiente y Fronteras: Una Visión desde los Países del Convenio Andrés Bello", pp. 79-91. Tercer Mundo Editores, Santafé de Bogotá.
  Marquet, P.A., L.C. Contreras, S. Silva & F.M. Jaksic (1993) Food habits of Pseudalopex foxes in the Atacama desert, pre-Andean ranges, and the high-Andean plateau of northernmost Chile. Mammalia 57: 130-135.
Marquet, P.A., L.C. Contreras, S. Silva, J.C. Torres-Mura & F. Bozinovic (1993) Natural history of Microcavia niata in the high Andean zone of northern Chile. Journal of Mammalogy 74: 136-140.
  Marquet, P.A., J. Pineda, J. Clark, M.J. Fortin, C. Jacobi, S. Wallins, J. Wu, S. Bollens & R.D. Holt (1993) Ecological and evolutionary consequences of patchiness. Lecture Notes in Biomathematics 96: 277-304.
Bozinovic, F. & P.A. Marquet (1991) Energetics and torpor in the Atacama Desert-dwelling rodent Phyllotis darwini rupestris. Journal of Mammalogy 72: 734-738.
  Jaksic, F.M., S. Silva, P.A. Marquet & L.C. Contreras (1991) Food habits of Gurney's buzzard in pre-Andean ranges and high Andean plateau of northernmost Chile. Journal of Raptor Research 25: 116-119.
Jiménez, J.E., P.A. Marquet, R.G. Medel & F.M. Jaksic (1991) Comparative ecology of Darwin's fox (Pseudalopex fulvipes) in mainland and island settings of southern Chile. Revista Chilena de Historia Natural 63: 177-186.
Jaksic, F.M, J.E. Jiménez, R.G. Medel & P.A. Marquet (1990) Habitat and diet of Darwin's fox (Pseudalopex fulvipes) on the Chilean mainland. Journal of Mammalogy 71: 246-248.
Marquet, P.A. (1990) Competition between distantly related taxa: three reasons why it is not more often reported. Revista Chilena de Historia Natural 63: 149-156.
  Marquet, P.A., F. Bozinovic, R.G. Medel, Y.L. Werner & F.M. Jaksic (1990) Ecology of Garthia gaudichaudi, a gecko endemic to the semiarid region of Chile. Journal of Herpetology 24: 431-434.
  Marquet, P.A., S.A. Navarrete & J.C. Castilla (1990) Scaling population density to body size in rocky intertidal communities. Science 250: 1125-1127.
Medel, R.G., P.A. Marquet, S.F. Fox & F.M. Jaksic (1990) Depredación sobre lagartijas en Chile central: importancia relativa de atributos ecológicos y morfológicos. Revista Chilena de Historia Natural 63: 261-266.
  Marquet, P.A. (1989) Paleobiogeography of South American cricetid rodents: a critique to Caviedes & Iriarte. Revista Chilena de Historia Natural 62: 193-197.
Marquet, P.A., J.C. Ortiz, F. Bozinovic & F.M. Jaksic (1989) Ecological aspects of thermoregulation at high altitudes: the case of Andean Liolaemus lizards in northern Chile. Oecologia 81: 16-20.
  Núñez, H., P.A. Marquet, R.G. Medel & F.M. Jaksic (1989) Niche relationships between two sympatric Liolaemus lizards in a fluctuating environment: The "lean" versus "feast" scenario. Journal of Herpetology 23: 22-28.
Medel, R.G., P.A. Marquet & F.M. Jaksic (1988) Microhabitat shifts of lizards under different contexts of sympatry: a case study with South American Liolaemus. Oecologia 76: 567-569.
  Ortiz, J.C. & P.A. Marquet (1987) Una nueva especie de lagarto altoandino: Lioleaemus islugensis (Reptilia-Iguanidae). Gayana Zoología 51: 59-63.